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Types of Receptors

   
There are three types of receptors in the body.  These receptors are constantly sending information back to the central nervous system about the success of a motor program or movement. 

The three types of receptors are:
1.  Skin Receptors
2.  Muscle Receptors
3.  Joint Receptors

The type of receptors we are most concerned with are muscle receptors.  

There are two types of muscle receptors: 
1.  Muscle Spindles
2.  Golgi Tendon Organs.

MUSCLE SPINDLES
Muscle spindles are located in the belly of the skeletal muscle, encased in a fusiform capsule.  A fusiform capsule is a capsule that is tapered on each end.  A rodlike structure runs through the capsule, and is divided into thirds.  The centermost third is the muscle spindle.  The two outer thirds are called intrafusal muscle fibers.   The entire capsule is encased in extrafusal muscle fibers.

While extrafusal muscle fibers (normal muscle fibers) are innervated efferently by alpha motor neurons, intrafusal muscle fibers are innervated efferently by gamma motor neurons.  

The muscle spindle is innervated by two different afferent sensory neurons.  
1.  Primary
2.  Secondary

Primary afferent neurons are sensitive to the amount and velocity of stretch placed on the muscle spindle.  Secondary afferent neurons are sensitive only to the amount of stretch.   

If the muscle itself is stretched, then the muscle will be stretched as well because the extrafusal and intrafusal muscle fibers are attached in series.  If the muscle spindle is stretched a threshold amount, then the primary and secondary afferent fibers will transmit a signal back to the central nervous system.  The CNS intreprets the signal, and understands that the amount of stretch in the muscle spindle could potentially cause structural damage to the muscle.  Therefore, a signal is sent down the alpha motor neuron to the skeletal muscle that encloses the muscle spindle, and the skeletal muscle contracts.  Ultimately, the stretch that could have been damaging to the skeletal muscle is prevented by contracting the muscle.  

GOLGI TENDON ORGANS

Golgi tendon organs are located where the tendon attaches to the muscle.  Unlike muscle spindles, they do not sense stretch - they sense excessive force and induce relaxation of the muscle via inhibition of the alpha motor neuron.  

 
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